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Melicope sandwicensis

(Hook. & Arn.) T.G.Hartley & B.C.Stone (1989)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Melicope

alani, alani kuahiwi

Description

Key Characters:

Growth Form: Small trees or shrubs 3–10 m tall, new growth densely minutely puberulent with tightly appressed, grayish hairs.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Blades narrowly elliptic to elliptic–ovate or elliptic–obovate, 4–15(–21) cm long, 2–5(–7.5) cm wide. Apex rounded to subtruncate, ± emarginate. Base rounded to subtruncate. Upper surfaces glabrous except midrib ± puberulent; lower surfaces sparsely to sometimes moderately puberulent, more densely so along midrib, ± pilose; blades subcoriaceous to chartaceous. Margins entire. Primary lateral veins usually 9–12 pairs, conspicuous, spreading-arcuate, connected by an arched vein 1–4 mm from margin, venation pinnate, lateral veins connected by an arched submarginal vein, higher order venation forming a reticulate pattern. Petioles 12–40 mm long. Stipules absent.

Flowers: Flowers 1–3(–6) in sparsely to densely puberulent, axillary cymes, peduncles (3–) 8–17 mm long, pedicels 3–8 mm long, bracteoles minute, caducous, bracts small, opposite. Calyx of 4 sepals; sepals oblong–ovate, 2.5–3.5 mm long, densely puberulent externally, glabrous within, margins ciliate, apex acute, subpersistent in fruit, imbricate. Corolla of 4 petals; petals oblong–deltate, 5–6 mm long, densely puberulent, apex obtuse, valvate. Stamens 8, in 2 whorls, distinct, reduced but always present in functionally pistillate flowers. Ovary superior, hirtellous; style ca. 0.4 mm long; stigmas clavate, not widely spreading. ovary 4-celled and 4-lobed, carpels connate, only slightly distinct at apex to almost entirely distinct; ovules 2 per cell, hemitropous, 1 ascending; style 1, terminating in 4 short stigmatic branches.

Fruit: Fruit probably dark green; 14–25(–30) mm wide; of distinct follicles 7–15 mm long; exocarp densely to moderately grayish or pale yellowish short–pilose to puberulent; endocarp sparsely to moderately short–villous with short; erect or appressed; pale hairs. the carpels dehiscing along the upper suture; exocarp. Seeds 1–2 per follicle; 4–7 mm long. glossy black when ripe; crustaceous; ovoid; sometimes angled from compression; embryo straight in fleshy endosperm; radicle short; cotyledons compressed; ovate; putamen brown; warty.

Ploidy:

Habitat: Occurring in diverse mesic forest.

Elevation Range: 450–850 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

O'ahu Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Taxon 38: 121 (1989)

Other References

Wagner et al. 1990:1204 (O [as Pelea sandwicensis]); Hartley & Stone 1989:121 (COMBNOV:Syn. P. sandwicensis)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Acacia koa-Metrosideros polymorpha lowland mesic forest PRESERVED_SPECIMEN Steve Perlman 10929 Oahu PTBG 4/11/1990
2 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Fosberg, F. R. 13016 Oahu US 3/22/1936
3 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN h. st. john 21512 Hawaii US 1/13/1946
4 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Fosberg, F. R. 13032 Oahu US 4/11/1936
5 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. montgomery & g. yamashita s.n. Oahu US 3/21/1971
6 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN St. John, H. 21512 Oahu US 1/13/1946
7 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Gifford, W. W. s.n. Hawaii US 8/9/2027
8 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Degener, O.; Greenwell, A. B.; Hatheway, W. 20683 Oahu US 7/4/1950
9 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN f. r. fosberg 13314 Oahu US 11/8/1936
10 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Montgomery, S.; Yamashita, G. s.n. Oahu US 3/21/1971
11 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Semi-moist wooded gully PRESERVED_SPECIMEN e. y. hosaka & o. degener 1938 Oahu US 1/13/1938
12 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN f. r. fosberg 13032 Oahu US 4/11/1936
13 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN f. r. fosberg 13016 Kauai US 3/22/1936
14 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Fosberg, F. R. 13314 Oahu US 11/8/1936
15 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN w. gifford s.n. Hawaii US 8/9/1927
16 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN g. pearsall s.n. Oahu US 7/27/1959
17 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Christophersen, E.; Wilder, G. P. 1611 Oahu US 2/20/1931
18 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Semi-moist wooded gully PRESERVED_SPECIMEN Hosaka, E. Y.; Degener, O. 1938 Oahu US 1/13/1938
19 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lay, G.T. Collie, A. Collector Number: s.n. Oahu BISH
20 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN russ, g.w. Oahu BISH 6/1/1930
21 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Pearsall, G. A. s.n. Oahu US 7/27/1959
22 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN e. christophersen & g. wilder 1611 Oahu US 2/20/1931
23 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN Gagne, W.C. 514 Oahu BISH 5/17/1970
24 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Eugenia sandwicensis, Metrosideros PRESERVED_SPECIMEN Nagata, K.M. 1169 Oahu BISH 7/4/1973
25 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Metrosideros, Straussia PRESERVED_SPECIMEN st.john, h. collector number: 25574 Oahu BISH 7/17/1955
26 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN o. degener, a. b. greenwell & w. hatheway 20683 Oahu US 7/4/1950
27 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 1776 o Oahu BISH 4/26/1912
28 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Takeuchi, W.N. 3490 Oahu BISH 2/1/1987
29 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information wet forest PRESERVED_SPECIMEN Cowan, R.S. 148 Oahu BISH 10/13/1946
30 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Christophersen, E. 1611 Oahu BISH 2/20/1931
31 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Ravine PRESERVED_SPECIMEN MacDaniels, L.H. 927 Oahu BISH 8/14/1927
32 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN cowan, r.s. collector number: 148 Kure Atoll BISH 10/13/1946
33 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information trailing over lantana PRESERVED_SPECIMEN st.john, h. collector number: 21512 Oahu BISH 1/13/1946
34 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Associated species: Psidium cattleianum, Grevillea robusta, Acacia koa, Schinus terebinthifolius, Metrosideros polymorpha, Psydrax odorata, Bidens torta, Clidemia hirta, Euphorbia multiformis, Alyxia stellata, Melinus minutiflora, Dianella sandwicensis. PRESERVED_SPECIMEN Ching, S. 20160630SCH01 Oahu BISH 6/30/2016
35 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Eugenia sandwicensis, Metrosideros PRESERVED_SPECIMEN nagata, k.m. collector number: 1169 Oahu BISH 7/4/1973
36 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information on Maba sandwicensis PRESERVED_SPECIMEN krauss, n.l.h. Maui BISH 12/30/1953
37 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN lay, g.t. collie, a. collector number: s.n. Oahu BISH
38 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN St. John, H. 21512 Oahu BISH 1/13/1946
39 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Ravine PRESERVED_SPECIMEN macdaniels, l.h. collector number: 927 Oahu BISH 8/14/1927
40 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN Degener, O. Takemoto, Martinez, M. Collector Number: 10580 Oahu BISH 4/21/1936
41 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN takeuchi, w.n. pyle, l.; tate, w.; lau, j. collector number: 2300 Oahu BISH 7/14/1985
42 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 1776 O Oahu BISH 4/26/1912
43 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information moist, wooded PRESERVED_SPECIMEN Hosaka, E.Y. 1938 Oahu BISH 1/13/1938
44 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information dry forest PRESERVED_SPECIMEN stone, b.c. collector number: 896 Oahu BISH 7/17/1955
45 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN degener, o. takemoto, martinez, m. collector number: 10580 Oahu BISH 4/21/1936
46 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN degener, o. collector number: 8523 Oahu BISH 7/10/1932
47 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Lay, G.T. s.n. Oahu BISH 12/30/1904
48 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information moist, wooded PRESERVED_SPECIMEN hosaka, e.y. degener, o. collector number: 1938 Oahu BISH 1/13/1938
49 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Metrosideros, Schinus, Planchonella PRESERVED_SPECIMEN Nagata, K.M. 1302 Oahu BISH 6/15/1976
50 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 1816 o Oahu BISH 4/26/1912
51 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3490 Oahu BISH 2/1/1987
52 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN gagné, w.c. collector number: 514 Oahu BISH 5/17/1970
53 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN Degener, O. 8523 Oahu BISH 7/10/1932
54 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN montgomery, s.l. yamashita, g. Maro Reef BISH 3/21/1971
55 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN christophersen, e. wilder, g.p. collector number: 1611 Oahu BISH 2/20/1931
56 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 1816 O Oahu BISH 4/26/1912
57 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information dry forest PRESERVED_SPECIMEN Stone, B.C. 896 Oahu BISH 7/17/1955
58 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information In pasture PRESERVED_SPECIMEN takeuchi, w.n. collector number: 3490 Hawaii BISH 2/1/1987
59 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Metrosideros, Schinus, Planchonella PRESERVED_SPECIMEN nagata, k.m. collector number: 1302 Oahu BISH 6/15/1976
60 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Krauss, N.L.H. s.n. Oahu BISH 12/30/1953
61 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN Degener, O. 10580 Oahu BISH 4/21/1936
62 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Metrosideros, Straussia PRESERVED_SPECIMEN St. John, H. 25574 Oahu BISH 7/17/1955
63 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN Degener, O. 11558 Oahu BISH 10/24/1937
64 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Russ, G.W. s.n. Oahu BISH 6/1/1930
65 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Takeuchi, W.N. 2300 Oahu BISH 7/14/1985
66 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN h. mann & w. t. brigham 558 Molokai F
67 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Mesic forest PRESERVED_SPECIMEN Steve Perlman 5793 Oahu PTBG 4/30/1987
68 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Steve Perlman 5276 Oahu PTBG 3/1/1987
69 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information Acacia koa- Metrosideros mesic forest. PRESERVED_SPECIMEN K. R. Wood 2924 Oahu PTBG 1/11/1994
70 Melicope sandwicensis (Hook. & Arn.) T.G.Hartley & B.C.Stone Contact Bishop Museum Botany department for locality information forest PRESERVED_SPECIMEN degener, o. salucop, f. collector number: 11558 Oahu BISH 10/24/1937
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